Paul Ullmann

GEOL 112 Discovering Dinosaurs! (*Satisfies Essential Studies Breadth of Knowledge - Math/Science/Technology requirement*)

GEOL 415 Introduction to Paleontology

GEOL 415L Introduction to Paleontology Lab

GEOL 416 Vertebrate Paleontology

GEOL 417 Dinosaur Paleontology

GEOL 418 Taphonomy and Fossilization Processes

GEOL 487 Research I

GEOL 488 Research II

GEOL 491 Geologic Problems

GEOL 494 Senior Thesis

GEOL 515 Advanced Paleontology

GEOL 591 Directed Studies

Taphonomy, Vertebrate Paleontology

PUBLICATIONS:

Gallucci JE, Woolslayer G, Barker K, Kibelstis B, Tumarkin-Deratzian AL, Ullmann PV, Grandstaff DE, Terry DO. 2024. Controls on soft tissue and cellular preservation in Late Eocene and Oligocene vertebrate fossils from the White River and Arikaree groups of Nebraska, South Dakota, and Wyoming. Minerals 14 (Special Issue: Recent Advances in Bone Diagenesis): 497. (invited paper) dx.doi.org/10.3390/min14050497

Boles ZM, Ullmann PV, Putnam I, Ford M, Deckhut J. 2024. New vertebrate microfossils expand the chondrichthyan and osteichthyan fauna of the Maastrichtian–Danian Hornerstown Formation in New Jersey. Acta Palaeontologia Polonica 69: 173–198. dx.doi.org/10.4202/app.01117.2023

Ullmann PV and Schweitzer MH. 2023. A statistical meta-analysis of lithologic and other potential controls on fossil bone cellular and soft tissue preservation. PALAIOS 38: 246–257. dx.doi.org/10.2110/palo.2022.026

Ullmann PV, Voegele KK, Lacovara KJ. 2023. Actualistic testing of the influence of groundwater chemistry on degradation of collagen I in bone. Minerals 13 (Special Issue: Recent Advances in Bone Diagenesis): 596. (invited paper) dx.doi.org/10.3390/min13050596

Ullmann PV, Ash RD, Scannella JB. 2022. Taphonomic and diagenetic pathways to protein preservation, part II: the case of Brachylophosaurus canadensis specimen MOR 2598. Biology 11 (Special Issue: Paleontology in the 21st Century): 1177. (invited paper) dx.doi.org/10.3390/biology11081177

Voegele KK, Ullmann PV, Boles ZM, Schroeter ER, Zheng W, Schweitzer MH, Lacovara KJ. 2022. Soft tissue and biomolecular preservation in vertebrate fossils from glauconitic, shallow marine sediments of the Hornerstown Formation, Edelman Fossil Park, New Jersey. Biology 11 (Special Issue: Paleontology in the 21st Century): 1161. (invited paper) dx.doi.org/10.3390/biology11081161

Schroeter ER, Ullmann PV, Zheng W, Schweitzer MH, Lacovara KJ. 2022. Soft-tissue, rare earth element, and molecular analyses of Dreadnoughtus schrani, an exceptionally complete titanosaur from Argentina. Biology 11 (Special Issue: Paleontology in the 21st Century): 1158. (invited paper) dx.doi.org/10.3390/biology11081158

Ullmann PV, Macauley K, Ash RD, Shoup B, Scannella JB. 2021. Taphonomic and diagenetic pathways to protein preservation, part I: the case of Tyrannosaurus rex specimen MOR 1125. Biology 10 (Special Issue: Paleontology in the 21st Century): 1193. (invited paper) dx.doi.org/10.3390/biology10111193  

Voegele KK, Ullmann PV, Lonsdorf T, Christman Z, Heierbacher M, Kibelstis BJ, Putnam I, Boles ZM, Walsh S, Lacovara KJ. 2021. Microstratigraphic analysis of fossil distribution in the lower Hornerstown and upper Navesink formations at the Edelman Fossil Park, NJ. Frontiers in Earth Science 9: 756655. dx.doi.org/10.3389/feart.2021.756655

Ullmann PV and Carr E. 2021. Catapleura Cope 1868 is Euclastes Cope 1868 (Testudines: Pan-Cheloniidae): synonymy revealed by a new specimen from New Jersey. Journal of Systematic Palaeontology 19: 491–517. dx.doi.org/10.1080/14772019.2021.1928306

McLain M, Ullmann PV, Ash RD, Bohnstedt K, Nelsen D, Clark R, Brand LR, Chadwick AV. 2021. Independent confirmation of fluvial reworking at a Lance Formation (Maastrichtian) bonebed by traditional and chemical taphonomic analyses. PALAIOS 36: 193–215. dx.doi.org/10.2110/palo.2020.064

Voegele KK, Ullmann PV, Lamanna MC, Lacovara KJ. 2021. Myological reconstruction of the pelvic girdle and hind limb of the giant titanosaurian sauropod dinosaur Dreadnoughtus schrani. Journal of Anatomy 238: 576–597. dx.doi.org/10.1111/joa.13334

Ullmann PV, Voegele KK, Grandstaff DE, Ash RD, Zheng W, Schroeter ER, Schweitzer MH, Lacovara KJ. 2020. Molecular tests support the viability of rare earth elements as proxies for fossil biomolecule preservation. Scientific Reports 10: 15566. dx.doi.org/10.1038/s41598-020-72648-6

Voegele KK, Ullmann PV, Lamanna MC, Lacovara KJ. 2020. Appendicular myological reconstruction of the forelimb of the giant titanosaurian sauropod dinosaur Dreadnoughtus schrani. Journal of Anatomy 237(1): 133–154. dx.doi.org/10.1111/joa.13176

Ullmann PV, Grandstaff DE, Ash RD, Lacovara KJ. 2020. Geochemical taphonomy of the Standing Rock Hadrosaur Site: exploring links between rare earth elements and cellular and soft tissue preservation. Geochimica et Cosmochimica Acta 269: 223–237. dx.doi.org/10.1016/j.gca.2019.10.030

Ullmann PV, Pandya SH, Nellermoe R. 2019. Patterns of soft tissue and cellular preservation in relation to fossil bone tissue structure and overburden depth at the Standing Rock Hadrosaur Site. Cretaceous Research 99: 1–13. dx.doi.org/10.1016/j.cretres.2019.02.012

Ullmann PV, Boles ZM, Knell MJ. 2018. Insights into cranial morphology and intraspecific variation from a new subadult specimen of the pan-cheloniid turtle Euclastes wielandi. PaleoBios 35: 1–22. escholarship.org/uc/item/8dw67415

Ullmann PV, Boles ZM, Lacovara KJ. 2018. Two new, large specimens of the pleurodiran sea turtle Taphrosphys sulcatus from Edelman Fossil Park, Mantua Township, New Jersey. The Mosasaur 10: 29–48.

Ullmann PV, Shaw A, Nellermoe R, Lacovara KJ. 2017. Taphonomy of the Standing Rock Hadrosaur Site, Corson County, South Dakota. PALAIOS 32(12): 779-796. (cover feature) dx.doi.org/10.2110/palo.2017.060

Ullmann PV, Bonnan MF, Lacovara KJ. 2017. Characterizing the evolution of wide-gauge features in stylopodial limb elements of titanosauriform sauropods via geometric morphometrics. The Anatomical Record 300(9): 1618-1635. dx.doi.org/10.1002/ar.23607

Ullmann PV and Lacovara KJ. 2016. Appendicular osteology of Dreadnoughtus schrani, a giant titanosaurian (Sauropoda, Titanosauria) from the Late Cretaceous of Patagonia, Argentina. Journal of Vertebrate Paleontology 36(6): e1225303. dx.doi.org/10.1080/02724634.2016.1225303

Lacovara KJ, Lamanna MC, Ibiricu LM, Poole JC, Schroeter ER, Ullmann PV, Voegele KK, Boles ZM, Carter AM, Fowler EK, Egerton VM, Moyer AE, Coughenour CL, Schein JP, Harris JD, Martínez RD, Novas FE. 2014. A gigantic, exceptionally complete titanosaurian dinosaur from southern Patagonia, Argentina. Scientific Reports 4: 6196. dx.doi.org/10.1038/srep06196

Ullmann PV, Varricchio DJ, Knell MJ. 2012. Taphonomy and taxonomy of a vertebrate microsite in the mid-Cretaceous (Albian–Cenomanian) Blackleaf Formation, southwest Montana. Historical Biology 24(3): 311-328. dx.doi.org/10.1080/08912963.2011.602405

PRESENTATIONS:

Kibelstis BJ, Terry DO, and Ullmann PV. 2023. X-ray diffraction analysis of fossil bones from the Paleogene White River Group of South Dakota and Nebraska: influence of apatite crystallinity on soft tissue preservation. GSA Connects 2023.

Armstrong Z and Ullmann PV. 2023. Actualistic assessment of early-diagenetic alteration of bones buried in highly-mature glauconitic greensands. GSA Connects 2023.

Boyd C, Drumheller S, Householder M, Saini-Eidukat B, Nestler JH, Ullmann PV. 2023. Geochemical examination of dinosaurian “mummies” from the Hell Creek Formation reveals a pathway for dermal tissue fossilization. Society of Vertebrate Paleontology Meeting Program & Abstracts.

Kibelstis BJ, Ullmann PV, Terry DO. 2023. Assessment of biomechanical function as a possible control on soft tissue preservation in Eocene–Oligocene bones from the White River Group of South Dakota and Nebraska. Geological Society of America Abstracts with Programs, Joint Northeastern/Southeastern Section Meeting: 20-43.

Ullmann PV. Braving the elements: taphonomic and diagenetic pathways to protein preservation. Keynote speech at the 2022 Annual Meeting of the Palaeontological Association, University College Cork, Ireland, July 2022.

Caputo C, Snyder K, Chadwick A, Ash RD, Ullmann PV. 2022. Using trace elements to characterize the geochemical history of the Hanson Ranch Bonebed, Cretaceous Lance Formation, Wyoming. Society of Vertebrate Paleontology Meeting Program & Abstracts: 99.

Woolslayer G, Ullmann PV, Grandstaff DE, Terry DO. 2022. The effects of taphonomic variability on the preservation of cells and soft tissues in fossil bones from Badlands National Park, SD. Geological Society of America Abstracts with Programs, Northeastern Section Meeting: 1-8.

Ullmann PV and Schweitzer MH. 2021. A statistical meta-analysis of lithologic and other controls on fossil bone cellular and soft tissue preservation. GSA Connects 2021: 2-2.

Voegele KK, Ullmann PV, Lonsdorf T, Christman Z, Lacovara KJ. 2021. Microstratigraphic analysis of fossil distribution in the lower Hornerstown and upper Navesink formations at the Edelman Fossil Park, NJ. GSA Connects 2021: 2-3.

Barker K, Terry DO, Ullmann PV. 2021. Recovering endogenous cells and soft tissues from fossil bones: does the depositional environment matter? GSA Connects 2021: 225-10.

Vieira J and Ullmann PV. 2021. Survivorship of invertebrate modes of life across the K/Pg boundary at Edelman Fossil Park, Mantua Township, New Jersey. GSA Connects 2021: 166-9.

Ullmann PV. 2021. Establishing criteria to differentiate mass-death and attritional fossil bonebeds based solely on trace element signatures. 9th International Bone Diagenesis Meeting: 39.

Ullmann PV. 2020. Why rare earth elements and other trace elements should be included in every vertebrate paleontologist’s research toolkit. TaphCon 2020: 40.

Vieira J, Voegele KK, Boles ZM, Ullmann PV. 2020. Building a comprehensive, multi-use fossil identification guide for the Edelman Fossil Park for education and outreach. GSA Connects Online 2020: 353979.

Grove J, Dallmann AL, Voegele KK, Ullmann PV, Nellermoe R. 2020. Contextualizing a new, large Allosaurus skeleton from Wyoming into stratigraphic perspective the Late Jurassic Morrison Formation. Geological Society of America Abstracts with Programs, North-Central Section Meeting: 16-15.

Macauley KW, Ash RD, Ullmann PV. 2020. Relating rare earth element uptake to soft tissue preservation in Cretaceous and Miocene fossil bones. Geological Society of America Abstracts with Programs, Joint Northeastern/Southeastern Section Meeting 52(2): 59-5.

Vieira J, Voegele KK, Boles ZM, Ullmann PV. 2020. Building a comprehensive, multi-use fossil identification guide for the Edelman Fossil Park for education and outreach. Geological Society of America Abstracts with Programs, Joint Northeastern/Southeastern Section Meeting 52(2): 9-6.

Engrav S, Voegele KK, Ullmann PV, Nellermoe R. 2020. Using Computed Tomography to analyze the braincase of a new Allosaurus specimen from the Jurassic Morrison Formation of Wyoming. Geological Society of America Abstracts with Programs, Joint Northeastern/Southeastern Section Meeting 52(2): 59-1.

Galluci J, Ullmann PV, Grandstaff DE, Ash R, Terry DO. 2020. Soft tissue and cellular preservation in Late Eocene-Early Oligocene vertebrate fossils of the White River Badlands. Geological Society of America Abstracts with Programs, Joint Northeastern/Southeastern Section Meeting 52(2): 40-7.

Voegele KK, Ullmann PV, Grove J, Nellermoe R. 2019. Characterizing the ontogenetic stage of a very large specimen of Allosaurus from the Jurassic of Wyoming with subadult characteristics. Society of Vertebrate Paleontology Meeting Program & Abstracts: 213.

Ullmann PV, Macauley K, Ash RD. 2019. Geochemical taphonomy of Tyrannosaurus rex MOR 1125, the first Cretaceous fossil to yield endogenous protein sequences. Society of Vertebrate Paleontology Meeting Program & Abstracts: 209.

Galluci J, Terry DO, Ullmann PV. 2019. Controls on soft tissue preservation in Eocene-Oligocene vertebrate fossils of the White River Group. Geological Society of America Abstracts with Programs 51(5): 211-2.

Galluci J, Ullmann PV, Terry DO. 2019. Soft tissue preservation in Late Eocene-Early Oligocene vertebrate fossils of the White River Group. Proceedings of the Eleventh Conference on Fossil Resources, Dakoterra 7.

Grove J, Voegele KK, Ullmann PV, Nellermoe R. 2019. Assessing the identify of a new Allosaurus skeleton from the Jurassic Morrison Formation of Wyoming. Geological Society of America Abstracts with Programs, Joint South-Central/North-Central/Rocky Mountain Section Meeting: 19-10.

Ullmann PV and Lacovara KJ. 2018. New insights into the osteology and phylogenetic affinities of the Cretaceous–Paleogene cheloniid sea turtle Catapleura repanda. Society of Vertebrate Paleontology Meeting Program & Abstracts: 231.

Boles ZM, Ullmann PV, Putnam I. 2018. Additions to the vertebrate fauna of Jean and Ric Edelman Fossil Park, including temporal range extensions across the K/Pg boundary. Society of Vertebrate Paleontology Meeting Program & Abstracts: 93.

Ullmann PV and Ash Rd. 2018. Characterizing the structure of diagenetically phosphatized K/Pg impact spherules from Edelman Fossil Park, Mantua Township, New Jersey. Lunar and Planetary Science Conference: 2767.

Ullmann PV and Lacovara KJ. 2017. Actualistic testing of the influence of groundwater chemistry on degradation of collagen I in bone. Geological Society of America Abstracts with Programs 49(7): 123-1.

Voegele KK, Ullmann PV, Heierbacher M, Kibelstis BJ, Putnam I, Lacovara KJ. 2017. Vertical distribution of fossils in the lower Hornerstown and upper Navesink formations at Edelman Fossil Park, NJ. Geological Society of America Abstracts with Programs 49(7): 220-12.

Ullmann PV and Lacovara KJ. 2017. Appendicular osteology of Dreadnoughtus schrani, a giant titanosaurian sauropod from the Late Cretaceous of Patagonia, Argentina. Society of Vertebrate Paleontology Meeting Program & Abstracts: 206.

Voegele KK, Ullmann PV, Boles ZM, Schroeter ER, Schweitzer MH, Lacovara KJ. 2017. Preservation of endogenous collagen I in a marine crocodile, Thoracosaurus neocesariensis. Society of Vertebrate Paleontology Meeting Program & Abstracts: 209.

Voegele KK, Ullmann PV, Lamanna MC, Lacovara KJ. 2016. Myological reconstructions from well-defined appendicular muscle scars in Dreadnoughtus schrani, a gigantic titanosaurian sauropod from Patagonia, Argentina. Society of Vertebrate Paleontology Meeting Program & Abstracts: 245.

Ullmann PV, Voegele KK, Grandstaff DE, Ash RD, Schroeter ER, Schweitzer MH. 2016. Evaluating the utility of rare earth element profiles as a proxy for soft tissue and biomolecular preservation potential in fossil bone. Society of Vertebrate Paleontology Meeting Program & Abstracts: 241. (invited talk)

Ullmann PV, Ash R, Grandstaff DE. 2016. Implications from diffusion modeling of trace element uptake by bones from the Standing Rock Hadrosaur Site, Hell Creek Formation, Corson County, SD. Geological Society of America Abstracts with Programs 48(7): 272-5.

Ullmann PV and Lacovara KJ. 2015. Evaluating the influence of body size on appendicular anatomy of titanosaurian sauropods. Society of Vertebrate Paleontology Meeting Program & Abstracts: 228.

Fowler EK, Voegele KK, Ullmann PV, Feldman V, Lacovara KJ. 2015. Restoring Dreadnoughtus: using lattice deformers in Autodesk Maya to retro-deform fossils from an exceptionally complete titanosaur. Society of Vertebrate Paleontology Meeting Program & Abstracts: 126.

Lacovara KJ, Lamanna MC, Ibiricu LM, Ullmann PV, Voegele KK, Schroeter ER, Boles ZM. 2014. An exceptionally complete titanosaurian sauropod dinosaur from the Upper Cretaceous of southern Patagonia, Argentina. Geological Society of America Abstracts with Programs 46(6): 264-14.

Ullmann PV, Grandstaff DE, Ash R, Lacovara KJ. 2014. Rapid and brief trace element uptake by bone at the Standing Rock Edmontosaurus bonebed, Hell Creek Formation, Corson County, SD: an exception to long-term rare earth element uptake. Geological Society of America Abstracts with Programs 46(6): 221-10.

Fowler EK, Ullmann PV, Voegele KK, Lacovara KJ. 2014. Titanosaur meets NextEngine: coaxing big data from an entry-level 3D laser scanner. Journal of Vertebrate Paleontology 34(5, Supp): 132.

Ullmann PV, Nellermoe R, Shaw A, Lacovara KJ. 2014. Taphonomy of the Standing Rock Edmontosaurus bonebed, Corson County, SD: implications for biomolecular preservation. Journal of Vertebrate Paleontology 34(5, Supp): 243.

Ullmann PV, Bonnan MF, Lacovara KJ. 2013. Morphometric exploration of the evolution of wide gauge features in stylopodial limb elements of titanosauriform sauropods. Journal of Vertebrate Paleontology 33(5, Supp): 229.

Voegele KK, Ullmann PV, Patel A, and Lacovara KJ. 2012. Insights from a new specimen of the gavialoid crocodilian Thoracosaurus neocesariensis from the Maastrichtian-Danian Hornerstown Formation, Sewell, NJ. Journal of Vertebrate Paleontology 32(5, Supp): 190.

Ullmann PV and Lacovara KJ. 2011. Largest known specimen and first mandible of the Cretaceous side-necked turtle Taphrosphys sulcatus (Testudines: Pleurodira). Journal of Vertebrate Paleontology 31(6, Supp): 207A-208A.

Ullmann PV, Varricchio DJ, Knell MJ, Lacovara KJ. 2010. Taphonomy and taxonomy of a vertebrate microsite in the Cretaceous Blackleaf Formation in southwest Montana. Journal of Vertebrate Paleontology 30(6, Supp): 179A.

2023                NSF GEOPAths Program (as CoPI):                                                            $179,992

2022                Open Access Publishing Fund, Rowan University:                                            $1,890

2021                Open Access Publishing Fund, Rowan University:                                            $1,721

2020                NSF MRI (for an ICPMS, as Senior Personnel):                                            $144,389

2020                Southern Adventist University, funds for a student researcher:                        $5,000

2018                Rowan University Seed Funding Program:                                                      $8,960

2014                Jurassic Foundation Grant:                                                                            $2,938

2010 - 2013      NSF Graduate Research Fellowship:                                                            $123,500

2009 - 2011      Provost’s Fellowship, Drexel University:                                                        $11,250

2008                 Top Senior in Geology Award, Department of Earth Sciences, Montana State University

2007 - 2008       Dr. C.C. Bradley Memorial Scholarship, Montana State University                        $600

2015 - 2017  Postdoctoral Scholar, Rowan University

2015  Ph.D. in Biology, Drexel University

2008  B.S. in Earth Sciences, Montana State University